Nettie Maria Stephens
Nettie Maria Stevens (1861-1912)
1905 discovered the X and Y sex chromosomes while working with beatles.
Stevens was able to provided proof for Calvin Bridges' hypothesis that genes were contained in chromosomes. She did this by examining the chromosomes of the fruit flys (Drosophila). Bridges wrote a couple of papers presenting the proof. He thanked her as "Miss Stevens" without stating what her contribution was nor refering to her PhD.