Starfish
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Starfish or sea stars are animals belonging to phylum Echinodermata, class Asteroidea. The name starfish is also used for the closely related brittle stars, which make up the class Ophiuroidea. They exhibit a superficially radial symmetry, typically with five or more "arms" protruding from a central body (pentaradial symmetry). In fact, their evolutionary ancestors are believed to have had bilateral symmetry, and sea stars do have some remnant of this body structure.
Starfish do not have movable skeletons, but instead possess a hydraulic water vascular system. The water vascular system has many projections called tube feet, on the ventral face of the sea star's arms, which function in locomotion and feeding.
Digestion
Starfish digestion is carried out in two separate stomachs, the cardiac stomach and the pyloric stomach. The cardiac stomach, which is a sacklike stomach located at the center of the body may be everted - pushed out of the organism's body and used to engulf and digest food. Some species take advantage of the great endurance of their water vascular systems to open the shells of molluscs (clams, mussels and the like), and inject their stomachs into the shells. Once the stomach is inserted inside the shell it digests the mollusk in place. Because of this ability to digest food outside of its body, the sea star is able to hunt prey that are much larger than its mouth would otherwise allow (including mollusks, arthropods, and even small fish). Partially-digested food is passed to the inside of the starfish where digestion continues in the pyloric stomach. Due to all of this digestive demand, the sea star's arms are filled with digestive glands called pyloric caeca or hepatic caeca...
External Anatomy
Sea stars are composed of a central disc with five arms exhibiting pentaradial symmetry. The mouth is located underneath (oral or ventral) the sea star, and the spiney surface covering the species is called the aboral or dorsal surface. On the aboral surface there is a structure called the madreporite which acts as a water filter and supplies their water vascular system with water to move (Gilbertson, 1999).
On the surface of the sea star surrounding the spines are small white objects known as pedicellariae. There consists thousands and thousands of these pedicellariae on the external body. Inside the sea star underneath all the hepatic caeca consist the gonads which are involved in reproduction. The radial canal which is across each arm of the sea star has what are called ampullae which surround the radial canal. The ampullae are teeth like structures.
Sea stars are developmentally (embryologically) known as deuterostomes. Since echinoderms and chordates share this same embryological pattern, they are thought to be closely related. Nevertheless, as these creatures are invertebrates and not actually fish, most marine biologists are pushing to completely replace the term starfish with sea star.
Regeneration
Some species of starfish have the ability to regenerate lost arms. When a starfish is cut into pieces, each part that includes a portion of the central disk may grow into a whole organism.
One genus particularly noted for its regeneration ability is Linckia, named for naturalist J.H. Linck. These starfishes can cast off an arm that regrows into an entire organism, as a means of asexual reproduction.
Geological history
Fossil starfish (Asteroidea) have been found in rocks as old as the middle Ordovician period. Brittle stars (Ophiuroidea) are first recorded as fossils in rocks from the Carboniferous period. Complete fossil starfish are very rare, but where they do occur they may be abundant. Most fossil starfish consist of scattered individual plates or segments of arms. This is because the skeleton is not rigid, as in the case of echinoids (sea urchins), but is composed of many small plates (or ossicles) which quickly fall apart and are scattered after death and the decay of the soft parts of the creature. Scattered starfish ossicles are reasonably common in the Cretaceous Chalk Formation of England.
Three famous localities where complete fossil starfish are found are the Devonian Bundenbach slates of Bundenbach in Germany, the Jurassic lithographic Solnhofen limestone of Solnhofen in Germany, and the Jurassic 'Starfish bed' of the Middle Lias formation near Bridport, Dorset in England.
See also
- Asterias
- Ophiuroidea (Brittle stars).
References
- Gilbertson, Lance; Zoology Lab Manuel; McGraw Hill Companies, New York; ISBN 0-07-237716-X (fourth edition, 1999)
- Solomon, E.P ., Berg, L.R., Martin, D.W. 2002. Biology Sixth Edition.