Gorgonops

Gorgonops Temporal range: Late Permian (Wuchiapingian)
Gorgonops whaitsi attacking Titanosuchus ferox
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Synapsida
Order:	Therapsida
Suborder:	Gorgonopsia
Family:	Gorgonopsidae
Genus:	Gorgonops Owen, 1876

Gorgonops (Gorgon (the mythological creature) and -ops, from Greek meaning 'face') is the name given to an extinct genus of synapsid which lived about 260 million years ago, during the late Permian Period. It was a typical representative of the Gorgonopsia, the dominant predators of their day, which in the largest forms grew to over 4 meters long and had evolved canine teeth, as well as other traits associated with its mammalian descendants. Arguments have even been made for synapsids of its time being warm-blooded, though no strong evidence exists either way.

Gorgonops itself was a medium to large-sized representative of the group, with a skull length of 20 to 35 cm, depending on the species.

The holotype of the type species, Gorgonops torvus, was in 1876 one of the first therapsids described, by Richard Owen, who also coined the name "Dinosauria" on the basis of the first known dinosaur fossils. It was also used as the type for which Richard Lydekker described the family in 1890. Five years later, in 1895, Harry Govier Seeley used this genus to establish the group as a whole. In later years, a large number of further species and genera were designated, but some of these turned out to be synonyms.

In Popular Culture

It featured in Episode 3 of Walking with Monsters as the top predator, preying on Scutosaurus.

Species

The following list of species follows the list of species provided Sigogneau-Russell, 1989. The type species is listed first, then the others in alphabetical order.

Since the above work was written, the dating of the Karoo Basin (Beaufort Group) has been revised. According to Smith and Keyser 1995, Gorgonops is known from the Tropidostoma and most of the Cistecephalus Assemblage Zones, and this has been taken into account when compiling the following list.

Gorgonops torvus OWEN, 1876 The type species. The holotype is an incomplete and flattened skull found at Mildenhalls, Fort Beaufort, South Africa. A number of other specimens have been found since, all from the Tropidostoma and/or Cistecephalus Assemblage Zone(s). This was a medium-sized animal, with a skull about 20 cm in length. It is distinguished from other species by a longer snout, and other details of the bones of the skull. Originally considered primitive, it is actually (according to Sigogneau-Russell) a rather specialised member of the group.

Gorgonops longifrons (HAUGHTON 1915) A large specimen known from an incomplete and flattened skull about 35 cm long. Orbit larger and snout longer than G. whaitsi, from which it may have descended. Beaufort West, Tropidostoma/Cistecephalus Assemblage Zone. Synonyms: Gorgonognathus longifrons HAUGHTON 1915

Gorgonops whaitsi (BROOM, 1912) Larger than G. torvus, with the rear of the skull wider, and other details of proportion. Originally the type species of Scymnognathus. Despite being known from a large number of specimens from the Karoo Basin, Beaufort West (Tropidostoma/Cistecephalus Assemblage Zone), the species remains poorly known. Watson and Romer placed Gorgonops and Scymnognathus in two different families, while Sigogneau-Russell 1989 placed the two species in the same genus, and considers G. whaitsi a more primitive (less derived) form. Synonyms: Scymnognathus whaitsi BROOM, 1912

The following are more problematic species (either synonyms or uncertain placement):

Gorgonops? dixeyi (HAUGHTON, 1926) A large, incomplete and flattened skull, from Chiweta Beds, Nyassaland. Genetic placement uncertain. Probably Low Cistecephalus Assemblage Zone equivalent (= middle of the Wuchiapingian Stage). See Jacobs et al 2005 for more on this species discussion on its age. Synonyms: Chiwetasaurus dixeyi HAUGHTON, 1926

Gorgonops? kaiseri (BROILI & SCHROEDER, 1934). A large (estimated total length about 35 cm long), incomplete skull, with a high snout and narrower in the rear than other species, from the "High Tapinocephalus zone" (i.e. earlier than the other species, most probably Pristerognathus Assemblage Zone) Synonyms: Pachyrhinos kaiseri BROILI & SCHROEDER, 1934

Gorgonops eupachygnathus (WATSON, 1921) A flattened, incomplete, medium-sized skull, probably a juvenile of either G. torvus or G. whaitsi Synonyms: Scymnosuchus whaitsi WATSON, 1921, Scymnognathus whaitsi (WATSON, 1921), Leptotrachelus eupachygnathus WATSON, 1921, Leptotracheliscops eupachygnathus (WATSON, 1921)

References

• Sigogneau-Russell, D., 1989, "Theriodontia I - Phthinosuchia, Biarmosuchia, Eotitanosuchia, Gorgonopsia" Part 17 B I, Encyclopedia of Paleoherpetology, Gutsav Fischer Verlag, Stuttgart and New York
• Jacobs, L. L., Winkler, D. A., Newman, K. D., Gomani, E. M. & Deino, A., 2005, Therapsids from the Permian Chiweta Beds and the age of the Karoo Supergroup in Malawi. Palaeontologia Electronica. Vol. 8, #1, pp. 28A: 21-23 online
• Smith, R.H.M. and Keyser, A.W. 1995. Biostratigraphy of the Tropidostoma Assemblage Zone. Geological Survey of South Africa, South African Committee for Stratigraphy, Biostratigraphic Series, 1:18-22.